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Anthropoid ape materials in American collections.

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ANTHROPOID APE MATERIALS
I N AMERICAN COLLECTIONS
WILTON M. KROGMAN AND ADOLPH H. SCHULTZ
Western Reserve University, Cleveland, Ohio, and Johns Eopkins Univereity,
Baliimore, Maryland
Foreword
At the New Haven meeting of the American Association
of Physical Anthropologists, April 30, May 1 and 2, 1936, a
committee on an 'Anthropoid Catalogue,' consisting of A. H.
Schultz (chairman) and W. M. Krogman, was appointed,
charged with listing all anthropoid materials in the United
States. In pursuance of this assignment over 150 questionnaires, based on a n earlier form drawn up by Prof. J. H.
McGregor, were sent o u t Institutions were listed as given
in Cobb's ('33) report on human materials, and zoological
gardens were circularized as summarized in Doolittle's ( '32)
report. In addition some sources of material were learned
through cross references. In all 135 questionnaires were returned either filled out or stating that no pertinent material
was available. Each institution or zoological garden possessing anthropoid material is listed by name in the tables comprising the bulk of this report. It is safe to say that a 95%
coverage of material has been achieved.
It was agreed in the interests of comparability that three
simply defined age classes should be drawn up: Infantile =
'Cobb, W. M. Human materials in American institutions available for anthropological study. Am. J. Phys. Anthrop., vol. 17 (Suppl.), pp. 15-16, April-June,
1933.
'Doolittle, W. 0. Zoological parks and aquariums. Am. Assoc. Zool. Parka
and Aquariums, vol. 1, pp. 98-100, Rockford, Ill., 1932.
199
AMERIOAN JOUKNAL O F PHYSICAL ANTH&DPOIl)aY, V O L X I I V , NO. 2
OCTOBBR-DECEMBER, 1938
200
W. M. EROGMAN AND A. H. SCHULTZ
birth to completion of deciduous dentition ; Juvenile =permanent dentition incomplete ; and Adult =permanent dentition
complete. Accordingly all materials are tabulated in these
age classes.
Of the perhaps Hteen or even more different species of
recent anthropoid apes only the two of the genus Gorilla have
been considered separately in this catalogue. The rare
Mountain Gorilla (Gorilla beringeri, which should really
have been named G. beringei) has been identified in all collections. Of Chimpanzees and Orang-utans the species are
frequently unknown and are at best difficult to determine from
skeletal parts alone, particularly since we do not as yet
possess generally agreed upon definitions for all of the
species. To have listed the many species of the Hylobatidae
separately would have made this report far too long.
The Committee wishes to thank all Directors and Curators
who so courteously and efficiently reported on the material in
their care. It is hoped that this unified catalogue, in making
available the knowledge of source material, will to some extent repay their kindness.
It must be emphasized that this census of anthropoid
material naturally can represent only the status of our collections at the particular time at which the questionnaires
were filled in. Changes will occur not only by the addition
of newly acquired specimens, but also by the deaths of living
apes which, it is hoped, will be saved as preserved bodies, and
by complete dissections of the latter, transfering them from
the tank to jars and the skeleton box. Specimens are sometimes exchanged between different collections. Finally, it
has occurred repeatedly that ape skeletons had been wrongly
labelled, but are in time transferred to their correct genus.
These and still other reasons are producing changes in the
collections.
201
ANTHROPOID APE MATERIALS
Skelet om
I n tables 1 to 6 are listed the complete skeletons of Pan,
Gorilla gorilla, Gorilla beringeri, Pongo, Hylobates, and
Symphalangus. As originally tabulated the material here
listed was stated to be ‘with or without skull,’ but an analysis
of the reports submitted shows that, barring a few exceptions,
a ‘complete skeleton’ means skull plus axial and appendicular skeleton. The tables may be summarized as follows:
Anthropoid skeletons in American collections
I
ADULT
.-
INFANTILE
I
2m
32
m
Pan
Gorilla gorilla
Gorilla beringeri
Pongo
Hylobates
Svmphalanguus
~
1
32
13
70
106 ! 65
4 : 2
55
..
14 1
11
24
17
1 1 1
~
11
2
..
F o r Pan the principal sources of material are to be found
in the Department of Mammalogy, American Museum of
Natural History, and in the Laboratory of Anatomy, Western
Reserve University. For Gorilla gorilla the chief source is
at Western Reserve University. For Pongo the chief source
is in the Division of Mammals, United States National
Museum. For Hylobates the principal sources are Department of Mammalogy, American Museum of Natural History,
Museum of Comparative Zoology, Harvard University, and
Division of Mammals, United States National Museum.
Skulls
I n tables 7 to 12 are listed the skulls of Pan, Gorilla gorilla,
Gorilla beringeri, Pongo, Hylobates and Symphalangus. As
a general rule each skeleton previously listed includes a skull
202
W. M. KROGMAN AND A. H. SCHULTZ
in these tables. For example, in table 1there are listed fiftyfive adult male Chimpanzee skeletons; in table 7 there are
listed eighty-seven adult male Chimpanzee skulls. Of these
eighty-seven skulls fifty-five are the skulls of skeletons listed
in table 1and thirty-two have no skeletons. In tables 7 to 1.2
no attempt has been made to state whether or not the mandible
is present. These may be summarized as follows :
Antltropoid skulls in American coZZectwns
I
ADULT
1
JUVENILE
INFANTILE
0
2
-
-.
______
Pan
Gorilla gorilla
Gorilla beringeri
Pongo
Hylobates
Symphaf angus
:
B &
-87
247
6
72
178
8
111
139
2
68
159
%
22
2
31
24
4
47
2
25
45
1
33
1
10
28
1
24
..
13
9
1
.
7
1
3
5
.
1
7
..
..
1
3
7
4
..
..
For Pan the principal sources of material are Department
of Mammalogy, American Museum of Natural History;
Division of Mammals, United States National Museum, and
Laboratory of Anatomy, Western Reserve University. For
Gorilla gorilla the principal sources are Department of
Anthropology, American Museum of Natural History;
Museum of Comparative Zoology, Harvard University;
Division of Mammals, United States National Museum, and
Western Reserve University. For Pongo the principal source
is United States National Museum. For Hylobates the
principal sources are Department of Mammalogy, American
Museum of Natural History; Field Museum of Natural
History ; Museum of Comparative Zoology, Harvard University; Collection of A. H. Schultz, Johns Hopkins University, and the United States National Museum.
203
ANTHROPOID APE MATERIALS
Miscellaneous skeletal material
In table 13 are listed miscellancous skeletal material for
Pan and Gorilla gorilla and in table 14 for Pongo, Hylobates,
and Symphalangus. These tables are summarized as follows :
Additional anthropoid skeletal matcrial in American collections
Pan
Gorilla gorilla
Pongo
Hylobates
Symphalangus
12
30
5
8
5
14
5
5
19
19
8
5
6 1
2
..
3
3
3
3
..
2
2
1
1
1
..
..
..
4
3
1
.
..
. .
.
..
.. 1
I
..
..
..
..
,
i
I
3
..
2
..
..
In this catalogue it is obviously impossible to identify the
miscellaneous skeletal material in detail. The numbers above
refer to separate individuals. For example, there are more
or less incomplete remains of twelve adult males of Pan,
thirty adult males of Gorilla gorilla, and so on. Since by f a r
the bulk of the miscellaneous bones for Pan and Gorilla
gorilla are found at the Laboratory of Anatomy, Western
Reserve Uxiiversity, it can be accepted from direct knowledge
that most fragmentary skeletons comprise a t least the long
bones. I n addition there may be vertebrae, scapulae, pelvic
bones, carpals, tarsals, phalanges.
Livimg aozthropoids
In table 15 the living specimens of Pan, and in table 16
those of Gorilla gorilla, Gorilla berengeri, Pongo, Hylobates,
and Symphalangus are listed. Two points must be kept in
mind in referring to these tables: 1) they are correct as of
October-December, 1937 ; and 2) several Zoological Gardens
and other sources (privately owned collections) have not replied despite frequent requests. With these limiting factors
in mind the tables may be summarized as follows :
204
W. M. EROGMAN AND A. H. SCHULTZ
JUVENILE
____
Male
Female
____ ____
INFANTILE
ADULT
Pan
Gorilla gorilla
Gorilla beringeri
Pongo
Hylobates
Symphalangus
__
!
25
I
Idale
1
.
l
I
2"
5
7
.
.
/
13
2
-j
2
..
1
3
4
..
Male
..
2
3
..
..
Female
__--
~
17
..
i
..
I
.! I
..
.3'
::
Entire animals preserved
I n table 17 are listed entire specimens of Pan, Gorilla
gorilla and Gorilla beringeri, and in table 18 of Pongo, Hylobates, and Symphalangus. The specimens are preserved in
formalin or other fluids and unless otherwise specified have
not been dissected or in any way mutilated. For P a n the
principal sources are Department of Comparative Anatomy,
American Museum of Natural History; and Laboratory of
Physical Anthropology, Johns Hopkins University. For
205
ANTHROPOID APE MATERIALS
Pongo in this respect the American Museum of Natural
History is the best source, while for Hylobates the American
Museum of Natural History; Museum of Comparative
Zoology, Harvard University ; Laboratory of Physical Anthropology, Johns Hopkins University ; and Division of
Mammals, United States National Museum, are the principal
sources.
MISCELLANEOUS UNDISSECTED
Parts or viscera
In table 19 are listed miscellaneous undissected partstrunk or limbs-and viscera-vascular, digestive, respiratory,
reproductive-of Pan, Gorilla gorilla, Gorilla beringeri,
Pongo, and Hylobates. Noteworthy are the partly dissected
bodies of Pan at the Laboratory of Physical Anthropology,
Johns Hopkins University, the limbs of Pongo in the collection of A. H. Schultz, the hearts and aortic arches of all the
anthropoids in the Department of Anatomy, University of
Oklahoma, and the finger, palm and sole prints of Pan in the
Department of Anatomy, Tulane University. The Department of Comparative Anatomy, American Museum of
Natural History, is the sole source for this type of material
of Gorilla beringeri.
-
_
_
Pan (28)
Gorilla gorilla (20)
Gorilla beringeri (1)
Pongo (47)
Hglobatea (47)
Symphalangus (8)
"I:
ADULT
Female
;
_
..
..
,
14
'
21
i
1
- J W E N I L E -Female
-Male
9
4
1.YFAWTIL.I
31ale
3
1
..
..
..
1
13
15
5
8
..
3
1
e
a
l
I
..
;
4
..
,
e
.
I
-
.
..
~Female
..
..
..
1
..
*.
~
206
W. M. KROQMAN A N D
A. H. SCHULTZ
The number in brackets is the total number reported. Insofar as age and sex was not given in every instance the
numbers classified do not always agree with the total. For
example there are twenty-eight brains of Pan, yet only fifteen
are classified by age and sex. The principal collection is that
of the Division of Physical Anthropology, U. S. National
Museum.
Mounted specimens
I n table 2 1 mounted specimens of Pan, Gorilla gorilla, and
Pongo are listed. It was the original intention to include
here not only articulated skeletons but mounted groups as
well. Obviously the information obtained is inadequate, for
the first type of material was most often referred to complete
skeletons, the second to hides. This section cannot, therefore,
be depended upon.
Skins
Information obtained on this item is as yet so defective
that the data would be of no value. Besides which this is an
item of zoological rather than anthropological importance.
Casts
I n table 22 casts of Pan, Gorilla gorilla, Gorilla beringeri,
and in table 23 of Pongo, Hylobates, and Symphalangus are
given, classified as to cast of head or skull, brain or endocast,
arm or leg, hand or foot. In several instances the ‘busts’ or
‘heads’ listed are commercial casts of Chimpanzees and
Gorillas. The same is true of ‘hands’ and ‘feet.’ As a rule,
however, the materials are unique to the laboratories reporting them. Especially is this true of the Department of
Comparative Anatomy, American Museum of Natural
History, and of the Museum of Comparative Zoology,
Harvard University, for Gorilla gorilla and Gorilla beringeri ;
and for the collection of A. H. Schultz which is equally representative for all of the anthropoids.
ANTHROPOID APE MATERIALS
207
Locality
The origina1,questionnaire did not call for the locality of
the material listed. When the majority of the reports were
in, it was deemed necessary to include locality. Accordingly,
all major sources of material, as judged by the returns, were
recircularized to ascertain locality of the material previously
reported. I n the following paragraphs will be summarized
the extent to which native provenience only, is known.
Material f o r which locality is unknown, or which has been
received from zoological gardens, will not be listed.
The Academy of Natural Sciences of Philadelphia reports :
F o r Pan, three skulls ‘from Africa,’ two skeletons with hides
of adult females, one skeleton with hide of juvenile male from
the Gaboon District; one skeleton with hide (adult female)
from Eufala, Cameroons; and one skeleton with hide of adult
female from Belgian Congo, Putnam’s Camp, Epulu River,
Kibale-Ituri. For Gorilla gorilla, three skulls ‘from Africa’
and one skeleton from West Africa; one skeleton from
Gaboon Country, Africa; one skeleton of adult male from
Ogove River, West Africa; three brains and skulls of juvenile
males from Kangive, Ogove River, West Africa; one skeleton
with hide of adult male from Lokundje, South Cameroons;
one skull of adult male from French Equatorial Africa, Nola
Region, Middle Congo; one skeleton with hide of adult male
from Burundu, 22 miles northeast of Nola; and three
skeletons with hides of adult males from Aboghi, 40 miles
southwest of Nola. For Pongo, six skulls, one listed as
female, are from Borneo. For Hylobates, a skull with hide
of adult male is from Burma, 40 miles east of Bhanad; a skull
with hide of adult male from Peninsula, Siam, Falls of Trang;
and skull and hide of adult male, skeleton and hide of adult
male and female from North Siam, Doi Sonteb, Chieng Mai,
4000’.
AYEEPICAN JOUPNAL OF PHYSICAL ANTHIWFQLI)OY, V O L XXIV, NO. 2
208
W. M. IiROOMAN A N D A. H. SCHULTZ
The Department of Comparative Anatomy, American
Museum of Natural History reports for Pan, all material
from the French Cameroons. For Gorilla gorilla, nine skulls
are from the Sanga River region, Quesso, West Africa, the
balance from the French Cameroons. For Gorilla beringeri,
Akeley ’s material, all from Kivu volcanoes, Belgian Congo.
F o r Pongo, an adult male skeleton from British North
Borneo. For Hylobates, all material from northern Burma.
The De’partment of Mammalogy, American Museum of
Natural History, reports for Pan, three from West Africa;
seven from Metet, Cameroons; two from Lukolela District,
Congo; one from Gamangin, Congo; seven from Medje,
Congo; nine from Faradje, Congo; two from Aba, Congo;
four from Akenge, Congo; four from Niapu, Congo ; one from
North G a p , Congo; one from Avakubi, Congo; one from
Eastern Congo; two from Uganda; one from Gaboon. For
Gorilla gorilla, two from Gaboon; three from Nola, French
Equatorial Africa ; one from N’Guilila, French Equatorial
Africa; one from Upper Nasable River; two from Ebole
Bengon, Cameroons; one from Metet, Cameroons; two from
Sakboyeme, Cameroons. For Pongo, all material from
Borneo. For Hylobates, one from Kuchin, Sarawak; five from
Yunan district, China; one ‘from China;’ five from Changchang Pani, Assam; one from Nakawu, Siam; nine from
Lampha, Tenasserim; one from Urn Pang, Siam.
The Department of Physical Anthropology, American
Museum of Natural History, reports for Pan, two from
Gaboon; four from Luemme; two from Ogawe; three from
South Cameroons; five from the Cameroons; eight from
Saunaga. For Gorilla gorilla, one from Ogawe; one from
Luemme ; three from the Cameroons ; two from Zaunde ; seven
from the Gaboon; four from Saunaga. F o r Pongo, two from
Borneo. For Hylobates two from Borneo.
The Carnegie Museum of Pittsburgh lists the Pan material
froan British East Afrioa md Lolodorf, Cameroons; the
Gorilla gorilla, from the Valley of the Ogove River, French
ANTHROPOID APE MATERIALS
209
Equatorial Africa, and Metet, Cameroons, West Africa ; the
Hylobates from Sumatra.
The Field Museum of Natural History reports for Pan:
five (two male, two female, one unclassified adult) from
Kitunda, Belgian Congo; four (two male, two female) from
the French Congo; one female from Nigeria, Cameroons,
Buec; one male from Nigeria, Cameroons, Mt. Cameroon;
one from Gabon; one female from Nyong River, Gabon; three
(one male, one female, one unclassified adult) from Cameroons, Gabon, Nyong River. For Gorilla gorilla, one male
from the Belgian Congo; one male from Cameroons,
Frontier; two (one male, one unclassified adult) from Dume,
Dume River, Cameroons ; one male from Lomie, Lomie River,
Cameroons; one male from Ebolowa, Cameroons; two males
from Yaunde, Yaunde River, Cameroons; one from Gabon;
one male from Ogave River, Gabon; one male from Kitunda,
Belgian Congo. For Gorilla beringeri one male from
Virunga volcanoes, Kigezi, Uganda. F o r Pongo one male
from Borneo; five (three males, two females) from British
North Borneo, 8 miles west of Sandakan. F o r Hylobates
three (one male, one female, one unclassified adult) from
Sandakan, British North Borneo; one female from Pang
Kallahau River, Southeast Borneo ; eleven (five males, six
temales) from Ban Me Thuot, Annam, French Indo-China.
The Laboratory of Comparative Zoology, Harvard University, reports the Pan material principally from Efulau,
Sakbayeme, Lolodorf, French Cameroons ; the Gorilla gorilla
principally from Metet and Sakbayeme, French Cameroons ;
the Pongo material principally from Abai, Kinabatangan
River, British North Borneo; the Hylobates material primarily from Chiengmai Region, North Siam, and Kinabatangan River, British North Borneo.
The Laboratory of Physical Anthropology, Johns Hopkins
University, reports, for Pan, one from Spanish Guinea, two
from the Cameroons. For Gorilla gorilla, three from
Spanish Guinea, all others from the Cameroons. For Pongo,
one from Borneo; and f o r Hylobates one from Java.
210
W. M, RROQMAN AND A. H. SCHULTZ
The collection of A. H. Schultz, Johns Hopkins University,
contains of Pan three specimens from the Cameroons, one
from Tangmyika. For Gorilla gorilla, all from the
Cameroons. For Pongo, seven from Sumatra, the others
from Borneo. F o r Hylobates, four from Borneo, eight from
Java, ten from Siam, two from Sumatra; and embalmed
material from the Malay Peninsula. For Symphalangus, all
from Sumatra.
The Department of Anatomy, University of Iowa, lists
for Pan one skeleton, for Gorilla gorilla two female skeletons, one male skull, one female skull-all from Abong Mbong,
Cameroons, West Africa.
The University Museums, University of Iowa, reports for
Gorilla gorilla, three male skulls, one female, one juvenile
skeleton, all from the French Cameroons. For Hylobates. one
male and one female adult skeleton, and one infantile skeleton,
are listed, all from Siam.
The Division of Mammals, U. S. National Museum reports
specimens as follows: Pan, twenty-seven from the French
Congo, twenty-three from South Cameroons ; one from Congo
Free State; three from Uganda; one from Sierra Leone.
Gorilla gorilla, twenty-seven from the French Congo ; twentythree from South Cameroons; three from the Belgian Congo.
Pongo, seventy-three from Borneo ;twenty-five from Sumatra.
Hylobates, ten from Sumatra; eight from West Sumatra;
twenty-four from East Sumatra; twenty-three from Borneo ;
five from East Borneo; six each from Southeast and West
Borneo; four from Southwest. Borneo; nine from South Pagi
Island ; eleven from Lower Siam ; one from West Siam ; seven
from Southeast Siam; four from Java; three from French
Indo-China; two from Cochin China; three from Johore; four
from Burma; one each from Pahang and Assam. Symphalangus, six from Sumatra; nine from East Sumatra; three
from Borneo; five from Southwest Borneo; two from Java;
one each from French Indo-China, Lower Assam, Tenasserim, and Pahang.
211
ANTHROPOID APE MATERIALS
The Laboratory of Anatomy, Western Reserve University,
lists Pan, fifteen from the Gaboon; 149 from Abong Mbong
and Ebolawa, French Cameroons; five from French Congo;
one from Belgian Congo; three from Nigeria; four from
French West Africa. Gorilla gorilla, thirty-two from the
Gaboon; 322 from Abong Mbong and Ebolawa, French
Cameroons; ten from French West Africa; eighteen from
French Congo. Pongo, ten from Borneo, ten from Sumatra.
Hylobates, ten from Borneo; four from Sumatra; three from
Burma; one from Java.
The Peabody Museum of Natural History, Yale University,
reports for Pan five skeletons from the Gaboon, three skeletons from West Africa; for Gorilla gorilla, four skeletons and
one skull from the Gaboon; for Pongo, one skeleton from
Sudong, Borneo and one skull from Borneo; f o r Hylobates,
one skeleton from Baram River, Sarawak, and one skeleton
from Sokaboini, Java.
SUMMARY
The most important materials of anthropoid apes in the
collections of the United States are summarized in the following table :
I
-
Pan
Gorilla
Pongo
Hylobates
Symphalangus
TOTALS OF SPECIMENS OF ALL AOES AND
BOTH s9Jl-B
272
246
149
263
14
400
541
230
455
23
98
7
17
22
2
73
11
31
64
9
212
W. M. KROQMAN A N D A. H. SCHULTZ
The living, preserved and cranial specimens of known sex
and all ages are combined and summarized in the following
table, which shows the sex ratios for the different genera.
The great preponderance of males in Gorillas, Orang-utans
and Gibbons, is particularly noteworthy.
QENUS
1
Pan
Gorilla
Pongo
Hylobates
Symphalangus
TOTALS OF BPECIMEIYS OB AL&
AQES AND -OWN
(Skulls, living animals and preaerved bodies)
Males
Females
207
321
125
277
12
241
182
99
221
17
The detailed data follow.
6 :9
X=
= 5 : 100
86
x = 176
LT = 126
x = 125
X=
SEX
71
TABLE 1
Pan (troglodytes). CompEete skeletolts
-
ADULT
I
s
3
4
3
il
m
-
3mzNILIp
INFANTILa
-
-i
m
I
-
a
1
m
%$
F
__
I
academy of Natural Sciences 0.
Philadelphia
American Museum of Natura
Hutory, N. P., Dept. of
Comparative Anatomy
American Museum of Natura
History, N. Y., Dept. of
Mammalogy
American Museum of Natura
N. Y., Dept. of Physical
Anthropology
Buffalo Museum of Science
Carnegie Museum,
Pittsburgh, Pa.
Cleveland Museum of Natural
History, Ohio
Field Museum of Natural
History, Chicago, 111.
Harvard University, Museum o
Comparative Zoology
Harvard University,
Peabody Museum
Harvard University, Warren
Anatomical Museum
Johns Hopkins University,
Laboratory of Physical
Anthropology
Johns Hopkins University,
Collection of A. H. Schultz
Philadelphia Zoological
Gardens, Pa.
PubHc Museum, Milwaukee, Wi
Science Museum of St. Paul
Institute, Minnesota
Southwestern University MuseuI
Memphis, Tenn.
Stanford University, Calif.,
Dept. of Anatomy
University of Iowa,
Dept. of Anatomy.
University o f Michigan,
University Museums
University o f Oklahoma,
Dept. of Anatomy
U. S. National Museum,
Division of Mammals
Western Reserve University,
Cleveland, Ohio, Lab. Anaton
Wistar Institute of Anatomy ax
Biology, Philadelphia, Pa.
Yale University,
Pea body Museum
Totals
-
3
_.
g
-2?
a
.
B
M%
;a
-
F
.. ..
..
1
..
__
..
3
2
1
1
1
1
..
1
1
..
..
..
8
3
6
.6
11
5
2
..
..
,.
..
..
..
..
..
..
1
..
..
..
..
..
..
..
8
5
4
1
..
..
..
..
..
..
1
..
..
..
..
..
..
..
1
2
5
..
1
..
..
4
6
..
1
6
..
1
4
1
2
..
..
..
..
1
..
..
..
..
2
4
..
5
..
..
..
2
1
..
..
..
1
..
1
..
..
..
..
..
5
18
1
..
..
2
..
2
1
..
..
..
..
..
..
..
..
,.
10
..
2
4
..
4
1
..
..
..
..
..
..
1
..
..
..
..
..
..
1
..
..
..
..
..
..
..
1
..
..
..
..
..
..
..
..
..
..
..
*.
..
..
..
1
..
..
1
..
..
..
..
7
..
3
1
..
..
38
2
5
9
13
..
1
..
..
..
..
..
..
..
..
..
..
..
..
..
a ,
I .
*.
..
..
..
..
..
7
..
2 .. .. .. 2 2 .. ..
1 11
6
32
5
40
22 31 70 55
- - ,.
..
..
213
..
TABLE 2
Gorilla (gorilla). Complete skeletons
MFANTILE
JUVENIU
ADULT
_-
-
-4 -
-e
a
s
0
a
B
Academy of Natural Sciences
of Philadelphia
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
American Museum of Natural
History, N. Y., Dept. of
Physical Anthropology
Army Medical Museum,
Washington, D. C.
Carnegie Museum,
Pittsburgh, Pa.
Cornell University, School of
Medicine, Dept. of Anatomy,
Ithaca, N. Y.
Field Museum of Natural
History, Chicago, Ill.
Harvard University, Museum of
Comparative Zoology
Harvard University,
Peabody Museum
Johns Hopkins University,
Lab. of Physical Anthropolog)
Public Museum, Milwaukee, Wis
Stanford University, Calif .,
Dept. of Anatomy
University of California,
Museum Vertebrate Zoology
University of Chicago,
Dept. of Anthropology
University of Iowa,
University of Museums
University of Iowa,
Dept. of Anatomy
University of Michigan,
University Museums
University of Wisconsin,
Dept. of Anatomy
U. S. National Museum
Division of Mammals
Washington University,
St. Louis, Dept. of Anatomy
Western Reserve University,
Cleveland, Ohio, Laboratory 0:
Anatomy
Wistar Institute of Anatomy ant
Biology, Philadelphia, Pa.
Yale Universitv.
Peabody Museum
Totals
L
6
..
2
..
..
..
..
..
..
1
2
..
..
..
..
..
..
1
3
3
2
2
..
1
..
..
..
1
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
1
..
..
..
1
..
3
..
..
..
..
..
..
..
..
..
..1
..
..
..
..
..
..
1
..
..
..
..
..
..
-
2
..
..
1
..
9
..
11
7
..
2
5
1
..
..
..
..
..
1
..
..
1
..
..
2
..
..
..
..
..
..
..
1
1
..
..
..
..
,.
1
..
1
..
..
..
1
..
2
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
1
11
1
1
..
..
4
1
1
5
4
..
..
..
..
..
..
53
38
..
1
..
..
..
..
..
16
14
..
1
..
..
..
..
..
..
..
..
..
..
..
..
..
..
3
..
.. 1 .. .. .. ..
..
.. 1 .. .. .. __
.. .. .. 1
2
5
65 6
22 13
106 17 ..
214
21.5
ANTHROPOID APE MATERIALS
TABLE 3
Gorilla (beringeri) Complete skeletons
.
I
ADULT
JUVENILE
-AXTILE
-P ;
-.
0
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
Field Museum of Natural Historj
Chicago, Ill.
Harvard University, Museum of
Comparative Zoology
Johns Hopkins University,
Collection of A. H. Schultz
U. S. National Museum,
Division of Mammals
Totals
--
-
2
1
..
..
1
....
....
..
....
..
1
..
..
-
32
m
..
..
.. ..
-
4
TABLE 6
Symphalangus (siamang). Complete sEeTetons
-- ADm
JUVENILE
INFANTIIg
I
a
--
m
-
I*
9:
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
..
American Museum of Natural
History, N. P.,Dept. of
Mammalogy
..
1
1
Johns Hopkins University,
Collection of A. H. Schultz
1
-.
. . . . ..
U. 8. National Museum,
Division of Mammala
..
I .
5
4
-6
5
1
....
__-
1
1
..
..
_
.
..
-
TABLE 4
Pongo (simio). Complete skeletons
__
-I
-$4 -
a
s
-s -i
INB'ANTIW
--
JUVENIlrB
ADULT
-.
$
3
2
4
1
..
rf - zi
m- 3 .. .. ..
1
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
..
2
..
..
..
..
..
1
..
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
1
..
1
10
2
4
..
..
..
American Museum of Natural
History, N. Y., Dept. of
Physical Anthropology
..
1
..
..
..
..
..
..
..
Field Museum of Natural
History, Chicago, Ill.
4
3
..
1
..
..
..
1
..
Harvard University Museum of
Comparative Zoology
3
3
..
2
1
..
2
..
..
Harvard University,
Peabody Museum
2
1
..
1
..
..
..
..
..
Johns Hopkins University,
Lab. of Physical Anthropology
1
..
..
1
..
..
..
..
..
Johns Hopkins University,
Collection of A. H. Schultz
1
..
..
1
1
2
Philadelphia Zoological
Gardens,' Pa.
1
..
..
..
..
..
..
..
..
..
1
..
..
..
..
Stanford University, Calif.,
Dept. of Anatomy
1
..
..
..
..
..
..
..
..
..
University of California,
Museum of Vertebrate Zoology
1
..
..
..
1
..
..
..
..
University of Wisconsin,
Dept. of Anatomy
1
1
..
..
..
..
..
..
..
12
21
..
3
..
1
..
.. ..
..
..
1
..
..
..
..
..
..
Cleveland, O., Lab. of Anatom
4
5
..
6
1
1
1
..
Wistar Institute of Anatomy
and Biology, Philadelphia, Pa.
4
2
5
..
..
2
..
..
..
..
..
3
3
..
..
..
2
..
..
..
..
-
t
a
m
Fr
Academy of Natural Sciences
of Philadelphia
Public Museum, Wilwaukee, Wk
U. 8.National Museum,
Division of Mammals
Waahington University,
St. Louis, Dept. of Anatomy
Western Reserve Univeraity,
Yale University,
Peabody Museum
Zoological Society,
Houston Texas
Totals
1
.. -
a
m
82
Y2
_
.
-
15
45
-39
216
-
26
~
m
I
m
.. .. ..
..
.. .. .. .. ..
2
5 11
6
..
-
TABLE 5
Hylobates. Complete skeletons
INPANTILAX
JUVENILE
ADULT
I
_
-
a
_
.
-
4
4*
4
-i
.cI
IJ
.cI
4
IH
Y
!::
Y
I::
m
m
__
Academy of Natural Sciences
of Philadelphia
1
1
..
..
..
..
..
..
..
American Museum of Natural
History, N. Y., Dept. of
comparative Anatomy
..
1
..
..
..
..
..
..
..
American Museum of Natural
HiNtory, N. Y., Dept. of
Mammology
5
8
2
5
1
..
..
..
..
..
1
..
..
..
..
..
..
..
3
10
..
3
..
..
..
..
..
60
52
3
13
14
2
5
3
1
Harvard University,
Peabody Museum
1
..
5
..
..
..
..
..
..
Johns Hopkins Umversity,
Lab. of Physical Anthropolog
1
..
..
..
1
..
..
..
..
Johns Hopkina University,
Collection of A. H. Schultz
9
3
..
1
..
..
1
..
..
..
1
..
..
..
..
..
..
..
University of Iowa,
University Museums
1
1
..
..
..
..
..
..
1
University of Michigan,
University Museums
1
..
..
..
..
..
..
..
..
19
10
1
..
..
..
..
..
..
Weatern Reserve University,
Cleveland, O., Lab. of Anatom
2
1
..
2
1
..
..
..
..
Wistar Institute of Anatomy
and Biology, Philadelphia, P
2
..
..
.’.
..
..
..
..
..
Carnegie Museum,
Pittsburgh, Pa.
F’ield Museum of Natural
History, Cliicago, Ill.
Harvard University, Museum o
Comparative Zoology
University of California,
Museum of Vertebrate Zoologg
0.8. National Mnseum,
Division of Mammals
Yale University,
Peabody Museum
Totals
1
.. -
105
90
3
14
-
~
217
.. .. .. .. .. ..
2
3
6
2
17
-
_
.
24
_
.
_
.
TABLE 7
ADULT
6
..
..
9
..
1
1
11
4
..
2
1
American Museum of Natural History, N. Y,
Dept. of Physical Anthropology
Buffalo Museum of Science, N. P.
Carnegie Museum, Pittsburgh, Pa.
Case School of Applied Science, Dept. of Geology
Cleveland, 0.
..
2
9
..
..
6
6
2
..
8
4
2
1
1
Field Museum of Natural History, Chicago, Ill.
Harvard University, Museum of
Comparative Zoology
Harvard Univer&ty, Peabody Museum
Harvard University, Warren Anatomical Museuc
Johns Hopkins University, Lab. of
Physical Anthropology
3
..
..
Cleveland Museum of Natural History, 0.
..
9
..
..
..
..
1
..
3
7
1
..
..
5
..
..
..
..
..
1
9
7
3
..
52
m
*
Pc
..
2
1
..
Cleveland Clinic Foundation, Cleveland, 0.
..
..
..
..
..
..
9
1
..
JUVENILE
..
1
1
..
..
..
..
..
3
..
2
6
..
..
7
..
2
_
I
i4
0)
-
Charleston Museum, 8. C.
3
1
American Museum of Natural History, N. Y.
Dept. of Mammalogy
1
1
2
$5
m
5*
a
American Museum of Natural History, N. Y.
Dept. of Comparative Anatomy
~
1
3
-~
F.1
B
Q
-
__
Academy of Natural Sciences, Philadelphia, Pa.
__
B
m
__
Pan (troglodytes). Skulls
..
..
..
..
..
..
7
..
1
..
..
..
..
..
..
..
..
1
..
..
..
.,
..
1
..
2
..
82
rn
f*
a
..
..
..
..
..
..
..
..
..
..
..
..
..
..
2
..
..
INFANTILE
..
..
University of California, Museum of
Vertebrate Zoology
1
..
36
..
2
..
..
Washington university, St. Louis,
Dept. of Anatomy
Western Reserve University, Cleveland, O.,
Lab. of Anatomy
Wistar Institute of Anatomy and Biology,
Philadelphia, Pa.
Yale University, Peabody Museum
Zoological Society, Houston, Tex.
Zoological Socity, San Diego, Calif.
87
..
12
U. 8. National Museum, Division of Mammals
Zoological Society, Toledo, 0.
Totals
1
1
University of Oklahoma, Dept. of Anatomy
1
111
..
..
57
..
13
..
..
..
University of Michigan, University Museums
..
..
..
University of Iowa, Dept. of Anatomy
..
1
Stanford University, Calif., Dept. of Anatomy
University of Iowa, University Museums
..
Southwestern University Museum, Memphis, Tenn
..
..
..
1
1
1
..
..
1
Public Museum, Milwaukee, Wis.
San Diego Museum, Balboa Park, Calif.
3
Philadelphia Zoological Gardens, Pa.
a .
Science Museum of St. Paul Institute, Minn.
4
Johns Hopkins University, Collection of
A. H. Schultz
3
..
37
..
..
..
..
..
38
..
-
2
..
I
1
..
4
..
..
..
..
2
..
..
..
..
1
..
41
..
-
..
..
..
..
13
..
..
..
,.
..
..
1
..
..
..
..
..
2
..
..
..
1
..
5
..
1
..
..
..
..
..
..
1
..
57
..
..
..
5
1
21
1
..
..
..
1
3
..
..
1
..
___
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
*.
..
..
..
..
..
..
..
..
..
..
..
..
..
* .
..
..
..
..
..
..
..
..
..
..
..
..
25
..
..
..
..
13
..
..
IQ
+
W
rn
E
E
3s
;dm
P
..
8
1
0
rd
zl3
P
..
..
..
..
..
1
..
..
..
TABLE a
Gorilla (gorilla).
Skulls
S0X
not
;ated
male
dale
_.
Academy of Natural Sciences,
of Philadelphia
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
American Museum of Natural
History, N. Y., Dept. of
Physical Anthropology
Army Medical Museum,
Washington, D.C.
Carnegie Museum,
Pittsburgh, Pa.
Case School of Applied Scienct
Dept. of Geology, Cleveland, C
Field Museum of Natural
History, Chicago, Ill.
Harvard University, Museum
of Comparative Zoology
Harvard University,
Peabody Museum
Johns Hopkins University,
Lab. of Physical Anthropolog
Johns Hopkins University,
Collection of A. H. Schultz
Philadelphia Zoological
ffarden, Pa.
Public Museum, Milwaukee, Wi
San Diego Museum,
Balboa Park, Calif.
Stanford University, Calif.,
Dept. of Anatomy
University of California,
Museum of Vertebrate Zoofog
University of Chicago,
Dept. of Anthropology
University of Iowa,
University Museums
University of Iowa,
Dept. of Anatomy
University of Michigan,
University Museums
University of Oklahoma,
Dept. of Anatomy
University of Wisconsin
Dept. of Anatomy
U. 8. National Museum
Division of Mammals
Washington University, St. Lou
Dept. of Anatomy
Western Reserve University,
Cleveland, O., Lab. of Anaton
Wistar Institute of Anatomy
and Biology, Philadelphia, Pa
Yale University,
Peabody Museum
Totals
-
INPANTILE
- - .-
_-
-
lale
male
-
-
3ex
not
ated
[ale
wale
9eX
not
ated
-
-
7
..
4
3
.. ..
..
..
..
4
3
4
2
..
1
..
..
..
1
3
9
..
..
1
..
..
..
13
8
1
..
1
1
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
4
..
..
1
..
4
2
1
..
..
..
..
1
..
1
..
..
1
1
..
..
..
9
..
..
1
..
15
10
..
2
3
1
..
..
1
..
..
..
5
4
1
..
..
..
1
2
..
..
..
..
..
1
..
1
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
1
1
..
..
..
1
1
1
..
3
..
..
..
..
1
..
..
1
1
..
3
1
..
..
..
I
I
..
..
3
1
..
..
1
..
..
..
1
I
..
32
If
..
..
..
..
3
2
..
..
1
..
,.
..
..
..
135
81
..
27
22
22
1
..
1
-
2
__
47
1
..
2 ..
139
241
-
22
-
220
..
..
..
1
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
..
1
..
..
..
..
.I
1
..
..
..
..
..
..
..
..
..
..
4
.. .. .. ..
..
.. 3
.. .. 7
1
7
33
24
..
-
- - -
-
221
ANTHROPOID APE MATERIALS
TABLE 9
Gorilla (beringmi). 8kulk
-
WL.T
-
I
INFANTILE
=
5
3*
m
i
fu
m
2
G
-
:m3
-
s
--
2
..
....
Field Museum of Natural History,
Chicago, Ill.
..
..
....
Hamard University, Museum of
Comparative Zoology
..
..
....
Johns Hopkins University,
Collection of A. H. Schultz
..
..
....
..
2
-
-
..
....
-....
-
American Museum of Natural
History, N. Y.,Dept. of
Mammalogy
U. 8. National Museum,
Division
-
of Mammals
..
TABLE 12
-~
Symphalangus (siamang). Skulls
--
1
_-INFANTILE-
ADULT
-I
I
0
i i
O
m
- -i
Q
Q
2
4=
*
u
--
; 3 -
American Museum of Natural
History, N. Y.,Dept. of
Comparative Anatomy
..
....
.. .. ..
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
1
1
.. .. ..
Hamard University, Museum of
Comparative Zoology
1
....
..
Johns Hopkina University,
Collection of A. H. Schultz
U. 8. National Museum,
2
.. i ..
. .I . .
--
.. .. ..
Division of Mammals
TOWS
1
.. ..
1
1
1
..
.. .. ..
..
- - ..
L
TABLE 10
Pongo ( h i a ) . Skulls
INFANT=
JUVIENILJe
ADULT
_-
a
.c1
-3
.,
*
$2
82
m
m
z a d e m y of Natural Sciences
of Philadelphia
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
American Museum of Natural
History, N. Y., Dept. of
Physical Anthropology
Camegie Museum,
Pittsburgh, Pa.
Field Museum of Natural
History, Chicago, 111.
Harvard University, Museum o j
Comparative Zoology
Harvard University,
Peabody Museum
Johns Hopkins University,
Lab. of Physical Anthropology
Johns Hopkins University,
Collection of A. H. Schultz
Philadelphia Zoological
Gardens, Pa.
Public Museum, Milwaukee, Wis
San Diego Museum,
Balboa Park, Calif.
Stanford University, Calif
Dept. of Anatomy
University of California,
Museum of Vertebrate Zoolog:
University of Iowa,
University Museums
University of Michigan,
University Museums
University of Wisconsin,
Dept. of Anatomy
U. S. Natioiial Museum,
Division of Mammals
Washington University,
St. Louis, Dept. of Anatomy
Western Reserve University,
Cleveland, O., Lab. of Anatom:
Wistar Institute of Anatomy
and Biology, Philadelphia, P a
Yale University,
Peabody Museum
Zoological Society,
Houston Tex.
Zoological Society,
Toledo, 0.
Totals
3
f*
0
--
4
3
9
..
..
..
..
..
..
..
..
1
..
..
..
..
..
..
1
..
1
4
2
4
..
..
..
3
4
3
..
1
..
..
1
..
..
..
..
1
..
..
..
..
..
5
5
5
3
..
..
..
1
1
5
3
..
2
1
1
..
..
3
2
1
..
1
..
..
..
..
1
..
..
..
..
..
..
..
..
..
4
3
..
1
1
6
..
..
1
..
..
..
..
..
..
..
..
..
..
..
1
..
..
..
1
..
..
..
..
..
..
1
1
..
..
..
..
..
..
..
1'
..
..
*.
1
..
..
..
..
..
..
1
..
..
..
1
..
..
..
1
1
..
..
..
..
..
..
..
..
..
..
..
..
..
..
33
35
..
4
4
..
..
..
..
1
..
..
..
..
..
..
..
..
5
6
..
7
1
1
..
..
4
1
8
..
..
..
..
..
..
..
..
3
3
..
..
1
2
..
..
..
..
..
..
..
1 .. ..
.. 10
..
..
..
..
72
-
1
..
..
1
1
~
68
31
-
222
25
-
-
..
..
.. ..
.. __
._ 7
1
3
TABLE 11
Hylobates. SkuUs
- -
--
a
B
:
-s z rn?22
- - 2
0
-Academy of Natural Sciencea
fi
INFANTILI
JUVENILE
,u
-4 -zi
0
-
-3
-a
1
..
..
2*
a
E2
rn
-
0
2
*
-.
a
94
W
3
2
..
..
American Mu~eumof Natural
History, N. Y., Dept. of
Comparative Anatomy
1
1
1
..
..
..
..
..
..
American Museum of Natural
History, N. Y., Dept. of
Mammalogy
10
8
1
6
4
3
..
..
..
5
17
..
16
2
..
..
..
..
61
53
7
15
14
4
5
3
1
1
..
5
..
..
..
..
..
..
1
..
..
..
1
..
..
..
..
10
13
..
1
3
..
..
1
1
*.
..
..
..
..
..
*.
..
..
of Philadelphia
Field Museum of Natural
History, Chicago, Ill.
Harvard University, Museum of
Comparative Zoology
Harvard University,
Peabody Museum
Johns Hopkins University,
Lab. of Physical Anthropology
John6 Hopkins University,
Collection of A. H. Schultz
Public Museum, Milwaukee, Wis
San Diego Museum,
..
__
..
2
..
..
..
..
..
..
..
1
..
1
..
..
..
..
..
..
..
University of Iowa,
University Museums
1
1
..
..
..
..
..
..
1
University of Michigan,
University Museums
1
..
..
..
..
..
..
..
..
U. S. National Museum,
Division of Mammals
67
57
5
..
..
..
..
..
..
Washington University,
St. Louis, Dept. of Anatomy
..
..
1
..
..
..
..
..
..
5
1
..
5
1
..
..
1
..
..
..
..
Balboa Park, Calif.
University of California,
Museum of Vertebrate Zoolog3
Western Reserve University,
Cleveland, O., Lab. of Anatom:
Wistar Institute of Anatomy
and Biology, Philadelphia, Pa.
Yale University,
Peabodv Museum
Totals
I
4 1
I
9
..
4
..
..
4
159
-
2
223
AMER[OAN SOVENAL OF PHYSICAL ANTHROPOLOQY, VOL. XXIY, NO. 2
..
..
I
I
..
..
.. 5
3
- -
_c
..
4
-
224
W. M. KROQMAN AND A. H. SCHULTZ
TABLE 18
Hiscellaneous skeletat material
I
I
ADULT
I
J-NLLID
INPANPILl
Pan (troglodytes)
-
-
.-
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
..
..
1
..
..
..
..
..
2
Carnegie Museum,
Pittsburgh, Pa.
..
1
..
1
..
..
..
..
..
Harvard University, Museum of
Comparative Zoology
3
1
..
..
..
..
..
..
..
Johns Hopkins University,
Lab. of Physical Anthropolog:
1
1
..
2
1
..
..
..
..
..
..
..
..
..
3
..
..
..
..
..
..
..
..
..
..
..
1
8
10
15
..
1
1
..
..
..
..
..
..
3
..
..
..
..
..
..
..
1
14
..
..
U. 5. National Museum,
Division of Mammals
Washington University,
St. Louis, Dept. of Anatomy
Western Reserve University,
Cleveland, O., Lab. of Anatom:
Yale University,
Peabody Museum
Zoological Society, Toledo, 0.
Totals
13
..
Gorilla (gorilla)
American Museum of Natural
History, N. Y., Dept. of
Comparative Anatomy
Harvard University, Museum of
Comparative Zoology
University of Iowa,
University Museums
U. S. National Museum,
Division of Mammals
Yale University,
Peabody Museum
Totals
..
3
..
I
1:
1
'
2
I
~
..
..
-
..
..
..
..
I
*.
..
..
.. ! ..
..
3
..
,
..
I
..
!
..
..__ _- ..
. .30
5
..
..
.. ! ..
..
..
10
1
3
3
' ..
..
..
5
19 '
1
3
..
..
__
-
3 !
..... -.-
,
1-3-
.
..
..
..
225
ANTHROPOID APE MATERIALS
TABLE 14
-
Pongo (simia)
American Museum of
Natural
History, N. Y.,Dept. of
Comparative Anatomy
..
..
1
.. ..
American Museum of Natural
History, N. Y.,Dept. of
Mammalogy
..
1
..
..
.. .. .. ..
Carnegie Museum,
Pittsburgh, Pa.
..
..
..
1
.. .. .. .. ..
Harvard University, Museum o
Comparative Zoology
..
..
6
..
..
.. .. .. ..
Johm Hopkins University,
Collection of A. H. Schultz
2
1
..
2
1
,. .. .. ..
Western Reserve University,
Cleveland, 0.; Lab. of Anatom:
..
1
1
.. ..
Wistar Institute of Anatomy
and Biology, Philadelphia, Pa.
3
1
..
..
1
..
5
5
8
.. .. .. ..
.. ..
.. .. .. .. 2
..
..
3
1
1
- -
Zoological
- Society,
Toledo, 0.
Totals
American Museum of
Natural
Hist.orv. N. Y.. Derk. of
Mammalogy
r
-
Johns Hopkins University,
Collection of A. H. Schultz
..
..
15"
2
.. .. ..
..
-
. . . . ..
..
-
.. ..
.-
3
..
--
..
..
Hylobates and Symphalangus a
Harvard University, Museum of
Comparative Zoology
U. 8. National Museum,
Division of Mammals
Totals
..
1
(S)
13 (5 5)
'Hylobates, except where (S) is stated.
..
I
&S)
....
..
*.
. . . . ..
..
. ... . . . . ..
226
W. M. KROGMAN AND A. H. SCHULTZ
TABLE 15
Living asthropoids. Pan (troglodytes)
ADULT
Female
..
Brookside Zoo, Cleveland, 0.
Central Park Zoo, N. Y.
3
City Zoo, Houston, Tex.
2
..
2
..
..
..
Como Park Zoo, St. Paul, Minn.
Male
-
hTPbNTm
Female
Yale
emale
-
1
1
..
..
1
2
..
..
..
..
..
..
..
..
..
..
..
..
..
1
Detroit Zoological Park,
Royal Oak, Mich.
3
1
..
Duluth Zoological Gardens, Minn
1
1
..
..
..
Fleishhacher Zoo,
San Francisco, Cal.
21
1(12)
..
..
L(2)
..
Forest Park Zoo,
F o r t Worth, Tex.
1
1
..
..
..
Franklin Park Zoological
Gardens, Boston, Mass.
1
1
..
..
..
..
..
..
..
..
..
..
1(7:
1(5)
..
..
..
..
..
..
..
.*
..
..
..
..
..
..
..
..
..
..
..
..
..
;:t
..
Hershey Zoological Gardens, Pa
Johmi Hopkins University,
Lab. of Physical Anthropologj
1
Lincoln Park Zoo, Chicago, Ill.
2 { E
Madison Zoological Gardens, Wis
..
1
..
..
3
1
..
1(9)
Municipal Zoo, Syracuse, N. Y
1
1
New Orleans Zoological
Gardens, La.
1(llt)
2 ( 11
124
..
New Pork Zoological Park, N. Y
1
4
Toledo Zoo, Ohio
1
1
..
Washington Zoo, Milwaukee, Wit
Yale University,
Lab. of Primate Biology
Zoological Society,
San Diego, Cal.
Totals
_____
4
1
-~
25
1
3
Philadelphia Zoological
Gardens, Pa.
..
..
..
15
1
..
1
1
1
1
5
10
-
2
2
1 __ .. __
..
3
38
13
17
-
-
~
..
..
-
..
2
-
227
ANTHROPOID APE MATERIALS
New York Zoological Park, N. Y.
..
Philadelphia Zoological Gardens, Pa.
1
Zoological Society, San Diego, Cal.
Totals
Central Park Zoo,
1
..
..
..
..
1( G.b.)
..
..
..
2(G.b.)
lL/
'2'
10 ..
.. ..
Pongo (simia)
N. Y.
..
Fleishhaeher Zoo, San Francisco, Cal.
..
Hershey Zoological Gardens, Pa.
..
Lincoln Park Zoo, Chicago, Ill.
I..
National Zoological Park,
Washington, D. C.
..
..
..
..
..
l(2t)
Philadelphia Zoological Gardens, Pa.
1
1
Zoological Society, San Diego, Cal.
1
..
Johns Ropkins University,
Collection of A. H. Schultz
II . .
5
Totals
..
..
..
..
..
1
1
.. 1(2f) __..3
1
2
-
Hylobates and Symphalangus
Central Park Zoo, N. Y.
Madison Zoological Gardens, Wis.
National Zoological Park,
Washington, D. C.
I
..
I1 . .
1
Philadelphia Zoological Gardens, Pa.
2
i1
Zoological Society, San Diego, Cal.
I
-1.7-
Gorilla gorilla except where (G.b.) is specified.
Hylobates except where (S) is specified.
I,
i
.. 1
1(5)
New 'York Zoological Park, N. Y.
Totals
.. I ..
12
I
1
'
..
I-.
..
1
,
!
.*
..
..
..
..
I ..
a
Pc
a
-
O
P a n (troglodytes)
parati& Anatomy
Cleveland Clinic
Foundation, Cleveland, Ohio
Forest Park ZOO,
Ft. Worth, Texas
Harvard University
Museum of Compai
tive Zoology
Johns Hopkins Un
versity, Lab. of
Physical AnthroPologY
Johns Hopkins Un
versity, Collection
A. H. Schultz
Philadelphia Zoo10
cal Gardens, Pa.
Tulane University,
New Orleans, La.,
Dept. of Anatomy
University of Wisconsin, Dept. of
Anatomy
U. S. National
Museum, Division
Mammals
Western Reserve U
versity, Cleveland,
0.. Lab. of Anatoi
Totals
-
-
3
6
2
..
1
.. ..
..
1
7
I{
j
1
1- -1I
ah
Totals
nt
--
..
..
1
.. .. .. .. . . ..
,.
.. ..
2
..
..
..
.. .. . .
..
..
..
..
1
.. .. ..
.. ..
2
..
4
..
9
3
..
..
..
..
..
1
1
.. ..
1
.. .. . .
1
.. 1
.. ..
_. .. ..
.. ..
1
.. .. ..
.. ..
..
.. .. ..
.. ..
..
1
..
..
..
3
..
1
2
-
...
.. .....
..
__
9
4
..
2
2 ___
_
15
.. ..
..
1
_.
12
'
..
1
1
,.
1
,.
.. .. .. .. ..
1.
1 .. _-.. - 1 .. _-1 ..
13
3
3
1
7
1
..
- - - - - ...
-
I
3
..
..
. . !. .
.. ..
..
..
..
. . , .. ..
..
1
I
..
.. ..
..
..
..
Anatnmv'
..
.. - - .' ~..
3(1G.b.)
~~
r
- -
-
.. ..
3
Gorilla (gorilla)
American Museum of
Natural History,
Dept. of Compara:(G.b.)
tive Anatomy
Carnegie Museum,
,
Pittsburgh, Pa.
i ..
Johns Hopkins University, Collection o f ,
A. H. Schultz
u. s. National
Museum, Division of
Mammals
..
Western Reserve Uni-,
-
I
.. ..
..
(
5
5
d
m
1
l(G.b.)
..
~
Gorilla gorilla, except where (G.b.) is specified.
228
..
..
..
11..
..
.. ..
.....
..
..
- 3
..
..
1
.. ..
..
-+
..
._
..
'
- 1_ _ _ _..
1
..
1
Academy of Natural
Sciences, Phila., Pa.
American Museum of
Natural History, N. Y.
Dept. of Comparative
Anatomy
Field Museum of
Natural History,
(Jhicago, Ill.
Harvard University,
Museum of Comparative Zoology
Johns Hopkins University, Collection of
A. H. Schultz
University of Wisconsin
Dept. of Anatomy
- 1
.. ..
-
..
- -
-
- - -
1
.. .. ..
..
.. .. .. ..
1
2
..
..
.. .. .. ..
..
.. .. .. .. .. ..
1
.. .. .. ..
..
..
1
..
..
..
..
..
4
5
.. .. .. .. ..
.. .. .. ..
1
.. .. .. ..
.. .. .. .. .. ..
..
.. .. .. ..
.. .. .. ..
..
..
1
1
P LlY
.. ..
pithout
head)
.. .. 1
.. 1 ..
.. .. .. ..
(b(
..
1
Z (one
..
.. ..
U, 8. National Museum
2
Division of Mammals
1
Washington University
(auto1
St. Louis, Dept. of
sied)
Anatomy
Western Reserve University, Cleveland, O.,
Lab. of Anatomy
Wistar Institute of
Anatomy and Biology ,
_-Philadelphia, Pa.
9
Totals
-
..
sset Id)
.. .. .. .. ..
1
1
1
..
.. ..
1 .. .. ..
.. .. -.. .. .. __
.. .. -.... 1 2
1
3 3 5
1
1
5 .. .. -I a es
American Museum of
Natural History, N. Y.
nept. of Comparative
Anatomy
I
Harvard University,
Museum of Comparative Anatomy
Johns Hopkins University, Lab. of
Physical Anthropology
Johns Hopkins University, Collection of
A. H. Schultz
U.. 5. National Museux
Division of Mammals
Wistar Institute of
Anatomy and Biology,
-Philadelphia, Pa.
Totals
- -
5
4
2
..
1
..
1
..
1
..
2
3
~
7
..
16
_
L
9yn
American
Museum of
Natural History, N. Y.,
Dept. of Comparative
Anatomy
Division of Mammals
U. S. National Museum,
Totals
5
1
2
3
3
.. ..
.. 5
..
__
4
m
. I
.. 3
1
11
-
gus
..
..
-
. . .
..
.. ..
.. ..
..
3
- __
TABLE 19
Yiscellaneol~sundissected parts or viscera
HPI.OBATES
--
..
American Museum of I limbs, 2 eyee, 1 male dissected 1 eyes
1set ovaries and limbs and trunk,
Natural History,
1 hands and feet
N. Y.,Dept. of Com- uterus
of (Q.b.) ; genito-'
parative Anatomy
urinary tract of
Cleveland Clinic
Foundation, Cleveland, Ohio
....
....
Ieart, thyroid, adrenals, genitals of
adult male
reproductive
tract, female
and misc.
viscera
Harvard University,
Museum of Comparative Zoology
Harvard University,
Warren Anatomical
Museum
'ongue, larynx,
f ace-skin, ears
(one animal)
Johns Hopkins University, Lab. of
Physical AnthroPologY
?artly dissected
bodies 1 adult f e
male, 4 juvenile
male, 4 infantill
female
...
Johns Hopkins University, Collection of
A. H. Schultz
....
....
....
....
....
limbs of infante; 2 hands,
2 feet of juve
niles; 1hand oi
adult
....
gall bladder
1 duodenum
,
.9 hearts and
Philadelphia Zoological Gardens, Pa.
tracts of viscer 1 reproductive
en masse
system (genitalia)
Tulane University,
New Orelans, La.,
Dept. of Anatomy
17 sets finger,
aortic arches;
1 lung
1 gall bladder
1 duodenum
heart and aortic 10 hearts and
aortis arches
arch
Oklahoma University,
Dept. of Anatomy
I
1tract of viacerr
en masse
....
palm, sole
of Yale C c E Z
animals
....
....
Western Reserve Uni- . ileo-cecal
versity, Cleveland, O., junction
Lab. of Anatomy
Wistar Institute of
Anatomy and
Bioloev. Phila.. Pa.
....
....
....
Harvard University,
Peabody Museum
Loyola University,
Chicago, Ill., Dept.
of Anatomy
....
....
....
~
sets limbs
....
....
hearts and
aortic arches
....
...
1 ileo-cecal
junction
1 male heart and
lungs
....
The Academy of Natural Sciences of Philadelphia has two jars of nnidentifIed and miscel-
laneous viecars.
* Gorilla gorilla unless (G.b.) specified.
230
TABLE 20
Brains
EYLOBATES
PAN
QOEXLL.4 1
PONQO
AND
SYMPEALANQUS
....
American Museum of Three brains
Natural History,
N. Y., Dept. of Com.
parative Anatomy
3ne brain
Academy of Natural Fwo brains
Sciences of Phila., P a
rhree braina
...
....
Cleveland Clinic
Foundation, Cleveland, Ohio
h e brain of adul rhree brains
female
....
....
Cornell University,
School of Medicine,
Dept. of Anatomy,
Ithaca, N. Y.
Fwo brains, one 3ne brain
left side (youni [young male)
males) ;one
brain, o m right
side (young
female)
Cwo brains
(adult female)
two brains, on
cerebellum
young female)
one brain (infantile female)
Harvard University,
Warren Anatomical
Museum
3ne-half of cerebrum
Philadelphia Zoological Gardens, Pa.
Fhree brains
University of Illinois, Phree brains (on'
Dept. of Anatomy,
hemisphere),
Ghicago, Ill.
mounted in
museum
U. S. National
Museum, Dept. of
Physical AnthroPOlOgY
3ix brains (5
young males, 1
young female)
Chree brains
....
....
...
Cwo brains
.,
h e brain (on
hemispere intact)
Fen brains (t
adult females
3 young males
1 young female), one
brain ((3.b.)
(young male)
....
....
'wo brains (H)
....
brains (5) (1
adult male, 2
adult females, 3
young males, 2
young females)
43 brains (H)
(21 adult males,
15 adult females, 3 young
male., 4 young
females)
....
....
h e adult male
one adult femal
'wo braina (H )
* Gorilla gorilla except where G.b. (= Gorilla beringeri) stated.
231
....
'4brains (13
adult males, 11
adult, females, I
young males, 6
young females:
Western Reserve Uni- Fwo adult male;
Fwo brains
versity, Cleveland, O., two juvenile
Lab. of Anatomy
male; one infan
tile female; one
adult and one in
fantile (sex unknown)
Wiatar Institute of
Anatomy and
Biology, Phila., Pa.
....
232
W. M. KROGMAN AND A. H. SCHULTZ
TABLE 21
Mounted specimens
PAN
I
PONQO
Carnegie Museum, Pittsburgh, Pa.
5, in group'
....
University of Iowa,
University Museums
1 juvenile male
Western Reserve University,
Cleveland, O., Lab. of Anatomy
1 adult female, 1 1 adult male 1 aduIt male, 1
Zoological Society, Houston, Tex.
'Listed under hides.
1
adult female,
1 juveniie
adult female,
1 juvenile
male
juvenile male, 1
adult .ight foot,
1 adult left
hand
....
L adult male,'
....
1 adult female
TABLE 22
casts
I
PAN
He+d, sknll,
brain, endocsst
Academy of Natural
Sciences, Phila., Pa.
American Museum of
Natural History, N.
Y., Dept. of Comparative Anatomy
Carnegie Museum,
Pittsburgh, Pa.
Field Museum of
Natural History,
Chicago, Ill.
Harvard University,
Museum of Comparative Zoology
Harvard University,
Peabody Museum
Harvard University,
Warren Anatomical
Museum
Johns Hopkins University, Collection of
A. H. Schultz
male head,
male busts
....
Limb, hand.
foot
....
....
5 heads
I.
....
....
skulls, 1 brain
hand, 1 foot
...
....
3 skulls, 1 brain 3 femora, 1
humerus, 1 tibia,
2 radii
1 face mask
1 hand, 1 foot
....
4 face masks
busts
.....
adult male
heads, 4 juvenil
male heads
adult male
)rain; 1 adult
male head; 2
adult male ears
Rochester Museum Art . adult male
brain
and. Sciences, N. Y.
. adult male
endocast
1 adult female
endocast
adult male sku
(G.b.)
(G.b.)
2 hands, 1 left
(G.b.); 1 head, foot (Q.b.), 2
1 adult male
left hands, 1
right foot
bust; 1 small
model entire
animal
....
hand
....
adult male an Heads: 2 adult Feet: 5 adult
male, 2 juvenile ( 2 are G.b.),
hands; 3 adult male, 1 juvenile 2 juvenile, 1 inmale and 4
female, 1 infan- fantile, 1foetus.
juvenile feet
Hands: 5 adult
tile female
(3 are Q-b.),
1 juvenile, 1 infantile, 1 foetus
adult male
1adult male brain
arms, 2 adult
male hands an
feet
adult male lei 2 adult male
skulls
foot; 1 adult
male right an
left hand
1 adult male
....
endoeast
4 juvenile
....
I
....
L endocast
Washington Universitj 1 brain, 1 skull
St. Louis, Dept. of
Anatomy
Western Reserve Uni- 1 adult male
versity, Cleveland, 0 head; 1 adult
female head
Lab. of Anatomy
-
1
male long bones 8 heads; 2 heads
(G.b.); 1 skull;
1 female and 2
male busts
brain
Public Museum,
Milwaukee, Wis.
San Diego Museum,
Balboa Park, Calif.
Science Museum of 81
Paul Institute, Minn.
University of Chicagc
Dept. of Anthropolog;
U. S. National Museum
Division of Mammals
Yale University,
Peabady Museum
....
heads
5 male and fe-
GO~LLA
1
Head, hll.
brain, endo-
Limb, hand
* Gorilla gorilla except where stated G.b.
....
....
....
....
....
2 adult male
1 skull, 1 brain,
1 endoeast; one,
small model entire animal
2 face-masks
....
....
....
5 right and left
adult male; 1 hands and feet,
face-mask adult 1 left foot
female; 3 endo-/ (G.b.)
casts
!
(= Gorilla beringeri)
.
234
W. M. KROGMAN AND A. H. SCHULTZ
TABLE 23
Costs
I
PONQO
Limb, hand,
foot
Academy of Natural 1 head
Sciences, Phila., Pa.
Limb hand.
fAot
....
....
....
....
....
American Museum of
Natural History, N.
Y., Dept. of a m parative Anatomy
2 heads
hands and feel
Carnegie Museum,
Pittsburgh, Pa.
1 brain
....
brain
....
Field Museum of
Natural History,
Chicago, Ill.
2 skulls; 1 brain
....
skull, 1 brain
....
Harvard University,
Museum of Comparative Zoology
2 adult male a m ,
1adult female skull
hand, 1 foot'
Harvard University,
Peabody Muse&
2 young heads
right hand
Johns Hopkins Uni-
2 adult male heads, Ianda and feet
adult head, 4
2 adult female
of 2 adult malei foetal heads
heads, 2 juvenile
male heads, 2 juve.
versity, Collection of
A. H. ESchultz
head
....
1 hand and
(a)
....
foot, 1
hand and
foot (a)
....
nile female heads
1 newborn head, 2
foetal heads
Public Museum,
Milwaukee, Wis.
Rochester Museum Arts
and Sciencks, N. Y.
San Diego Museum,
Balboa Park, Calif.
....
....
....
....
....
adult male sku1
....
1 adult male
endocast
....
1 adult male brain
University of Chicago,l adult male skull
Dept. of Anthropology
Yale University,
Peabody Museum
1 brain, 1 endocast
Western Reserve Uni- 2 adult male heads
versity, Cleveland, O.,
Lab. of Anatomy
adult female
endocast
....
....
....
....
....
....
....
....
endocast
....
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